Half of all insect species are dependent on living plant tissues, consuming about 10 per cent of plant annual production in natural habitats and an even greater percentage in agricultural systems, despite sophisticated control measures. Plants possess defences that are effective against almost all herbivorous insect species. Host-plant specialization, observed in over 80 percent of these animals, appears to be an effective adaptation to breach these defence systems. The mechanisms underlying plant defence to invading herbivores on the one side, and insect adaptations to utilize plants for...