Analytic philosophers once pantomimed physics: they tried to understand the world by breaking it down into the smallest possible bits. Thinkers from the Darwinian sciences now pose alternatives to this simplistic reductionism.

In this intellectual tour--essays spanning thirty years--William Wimsatt argues that scientists seek to atomize phenomena only when necessary in the search to understand how entities, events, and processes articulate at different levels. Evolution forms the natural world not as Laplace's all-seeing demon but as a backwoods mechanic fixing and re-fashioning...