Heliconiinae is one subfamily of Nymphalidae, has played a key role in understanding evolutionary biology and ecology and it would be difficult to point a group of Neotropical butterflies that have contributed more to our knowledge of the biological processes in the tropics. The derived members of the tribe have undergone rapid speciation and divergence, while also exhibiting impressive mimetic convergence in wing patterns. Despite several recent important efforts to elucidate the higher level relationships of butterflies, there is still only fragmentary knowledge about patterns of...