The increasing interest during recent years in vegetation history, long-term climatic changes and the development of biotic communities has resulted in a growth of the literature in this field and has provided an incentive for scientists to use palaeo-ecology concepts to clarify contemporary ecological issues. Quaternary ecology offers the perspective of time, in using natural experiments of the past to evaluate the responses of population, communities and ecosystems to climatic and environmental changes on a time scale extending back over the past two million years of the quaternary period....