Quantitative models are crucial to almost every area of ecosystem science. They provide a logical structure that guides and informs empirical observations of ecosystem processes. They play a particularly crucial role in synthesizing and integrating our understanding of the immense diversity of ecosystem structure and function. Increasingly, models are being called on to predict the effects of human actions on natural ecosystems. Despite the widespread use of models, there exists intense debate within the field over a wide range of practical and philosophical issues pertaining to...