Population genetics thrives on the constant interaction between theoretical and empirical knowledge. In the first instance, population genetics was developed using a one locus, two allele model for genetic variation. The simplicity of these models opened up population and evolutionary genetics to biologists without specialist training in mathematics. Population genetics of multi-allelic loci is more complex and requires a degree of mathematical insight, therefore its study is In this presentation, the author provides a unified theory which demonstrates that multi-locus theory need not be an...