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"As a general principle natural selection is continually trying to economise every part of the organisation. " That was Charles Darwin, writing over 100 years ago about e- ciency innature. Naturalmaterialsareremarkably ef?cient. Byef?cient wemeanthat they ful?l the complex requirements posed by the way plants and animals function, and that they do so using as little material as possible. Many of these requirements are mechanical in nature: the need to support static and dynamic loads created by the mass of the organism or by wind loading, the need to store and release elastic energy, the need...