1. Introduction; 2. Getting Started – Data and Phylogeny; 3. Phylogenetic Non-independence; 4. Tempo and Mode – Brownian Motion and More; 5. Incorporating Estimates of Error; 6. Variation Across Trees; 7. Model Fit and Model Adequacy; 8. Post-hoc Modelling of Discrete Characters; 9. Modelling Heterogeneous Trait Evolution; 10. Conclusions; References.