Richard Darwin Keynes is Emeritus Professor of Physiology at the University of Cambridge, and fellow of Churchill College, Cambridge. In parallel with research and teaching on physiology and biophysics, he has a long-standing interest in the voyage of the Beagle, and has also edited The Beagle Record (1979) and Charles Darwin's Zoology Notes and Specimen Lists from the H. M. S. Beagle (2000).