Born 1934, studied at the University of Zurich (General Botany, 1970 Dr. phil. II). 1967-1986 embryological and cytological analyses of progeny of sexual x apomictic biotypes of Ranunculus auricomus (up to BC3). Habilitation ETH Zurich 1978. editorial work. 1987-1999 Archivist at the University of Zurich.